DOI: 10.3897/pls2024612/01

Authors: Silvia Assini, Ilaria Brugellis, Juri Nascimbene, Matteo Barcella, Alessia Gressani, Gabriele Gheza

Abstract: This paper provides an overall look on the diversity of lowland dry grasslands of the western Po Plain, useful to support their conservation and management. Specific aims were: 1) to identify lowland dry grassland-associated plant communities at alliance level, which is useful for their classification under the Council Directive 92/43/EEC, 2) to assess their synecological and synchorological differences, 3) to define the equivalent Directive habitats and their management implications. Seven subareas were analysed: Lomellina, Ticino River, Sesia River, Po River (North), Orba Stream, Scrivia Stream, and Serio River. Floristic-vegetational relevés were carried out considering vascular plant, moss and lichen species. Cluster analysis were performed to syntaxonomically classify them, while statistical tests were performed to characterize them by means of biological life forms, chorotypes and Ellenberg indicator values. Eight plant communities were classified at alliance level and three plant communities were classified at class level. The equivalence with three Natura 2000 Habitats (H2330, H6110* and H6210) was found. Of the 60 studied sites, the 68% are located inside the Natura 2000 Network, while the remnant 31% are located outside. Possible management actions include: cutting of woody species, mowing, sod-cutting, transplants of typical herbaceous species, and ex novo restoration using harvested seeds from donor grasslands.

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DOI: 10.3897/pls2023602/01

Authors: Erwan Glemarec, Frédéric Bioret

Abstract: Atlantic coastal cliff heathlands are chamaephytic vegetation dominated by Ericaceae, most often associated to Fabaceae (Ulex) and integrated from a phytosociological point of view into alliance of Dactylido oceanicae-Ulicion maritimi Géhu 1975 (class of Calluno vulgaris-Ulicetea minoris Braun-Blanq. & Tüxen ex Klika & Hadač 1944). This alliance gathers coastal heathlands under marine influences whose physiognomy is marked by prostrate or halo-anemomorphic chamaephytes. All Atlantic heathlands are included into habitats of European community interest which justify the designation of Natura 2000 sites. Within these protected areas, heathland conservation or restoration management issues raise the question of the choice of nature and frequency of management methods. For that, it seems important to understand the dynamics of cliff-top coastal heathlands. Stable heathlands, considered as permanent vegetation of primary character, can be managed without any intervention, while heathlands characterized by a progressive dynamic may be subject to interventionist management. The aim of this work is to present the main determinant abiotic and biotic factors driving the cliff-top coastal heathlands. Human uses may influence the composition and structure of these vegetation. This approach helps to assess the primary or secondary character of cliff-top coastal heathlands vegetation, their stability or dynamic. This article clarifies the notions of primitive, primary and secondary coastal heathlands. It proposes criteria to differentiate stable heathlands and unstable heathlands which are characterized by a progressive dynamic towards coastal scrublands or forest: abiotic conditions, floristic combination, physiognomy. The synphytosociological methodology is useful for establishing the typology of the series, permaseries and minoriseries, in which the different types of cliff-top coastal heathlands are included.

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DOI: 10.3897/pls2023602/05

Authors: Danièle Lagnaz, Giacomo Trotta, Patrice Prunier, Bertil Krüsi, Francesco Boscutti

Abstract: This paper presents a phytosociological study of dry and mesophilous meadows and pastures in the Locarnese region (Insubria - Southern Switzerland). Seventy-one vegetation relevés were analysed using both hierarchical classification and nonmetric multidimensional scaling (NMDS) ordination. Seven main clusters were identified and described as follows: (i) xerophytic grasslands on sandy soil dominated by Koeleria macrantha; (ii) mesophilous meadows with Arrhenatherum elatius and Centaurea transalpina; (iii) semi-dry meadows with Chrysopogon gryllus; (iv) semi-dry grasslands with Carex fritschii and Thalictrum minus; (v) nutrient-poor Phyteuma betonicifolium-Festuca nigrescens montane zone grasslands; Nardus stricta grasslands dominated by (vi) Festuca paniculata or (vii) Carex pilulifera. From a phytosociological point of view, a new association named Phyteumo betonicifolii-Festucetum nigrescentis, classified into the Nardo strictae-Agrostion tenuis alliance was proposed. In addition, we confirmed the presence of the Holco-Chrysopogonetum grylli association, within the Bromion erecti, which up to now was provisional only.

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DOI: 10.3897/pls2023601/06

Authors: Gianmaria Bonari, Michele Dalle Fratte, Michele Lonati, Marco Caccianiga, Cesare Lasen, Stefano Armiraglio, Matteo Barcella, Gabriella Buffa, Bruno Enrico Leone Cerabolini, Andrea Mainetti, Luca Miserere, Giuseppe Oriolo, Alberto Selvaggi

Abstract: Habitats Directive (92/43/EEC) is the cornerstone of nature conservation in Europe and is at the core of the EU Biodiversity Strategy for 2030. There is room, however, for its improvement, at least for northern Italy, where ambiguities in the definition of habitat types of Annex I of the Habitats Directive are not novel and interpretation difficulties have been highlighted. Sharpening the characterization of habitat types represents an opportunity for lowering classification uncertainties and improving conservation success. With the aim to refine the definitions of habitat types and associated typical species of the Habitats Directive, a group of vegetation scientists of the Italian Society of Vegetation Science based in northern Italy made the exercise of finding viable proposals for those habitat types having a problematic interpretation in the Alpine biogeographical region of Italy. Such proposals arise from group discussions among scientists, and professionals, thus offering a shared view. We prepared 9 habitat proposals important for this geographic area. They include new habitat types at the European level, new subtypes within pre-existing habitat types, including some adjustments of the recently proposed subtypes with respect to northern Italy, and recognition of priority criteria for a pre-existing habitat type. With a vision of tailored conservation, our proposals represent a starting point in view of a future update of Annex I. Furthermore, the list of typical species could be useful for preparing expert systems for automatic classification. Irrespective of legally binding solutions in place, we caution these proposals represent relevant baseline conservation indications that local and regional administrations of the Alpine Arch should consider.

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DOI: 10.3897/pls2023601/04

Authors: Naim Berisha, Renata Ćušterevska, Fadil Millaku, Vlado Matevski

Abstract: The sedge-moss vegetation of the moderately to low calcium-rich slightly acidic fens of the Caricion fuscae alliance depends on a very specific combination of ecological and climatic conditions to thrive. Until recently, the classification of this vegetation group was complicated by its rarity on the southern edges of its range in Europe. As part of a larger database of phytocenological relevés carried out in Mt. Luboten, we came across an interesting group of 15 relevés on fen vegetation sites. We were curious to know if this plant community was a previously known association or if it might represent something new within this alliance. We compiled a separate dataset at JUICE that includes four plant communities from this alliance, along with our 15 original releves. The classification was based on modified TWINSPAN and beta-flexible clustering as a numerical classification method, with OPTIMCLASS determining the appropriate number of clusters. Five associations were clearly delineated, with the four associations taken from the literature sources clearly grouped individually and a new, fifth association appearing as separate, with completely unique characteristics. This new association: Blysmo compressi-Eriophoretum latifoliae occurs at elevations of ~ 1650 m a.s.l. on NE and NW slopes of the mountain. With this work we offer the description of a new high-mountain fen association. These associations may play an important syntaxonomic role as more Balkan data become available on this alliance. The sedge-moss and fen vegetation in the Balkans is particularly rare and characterised by a very diverse and specific vegetation, so it rightly deserves more attention from vegetation scientists and conservation authorities.

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DOI: 10.3897/pls2023601/01

Authors: Harrie G.J.M. van der Hagen, Erik Lammers, Frank van der Meulen, Ricarda Pätsch, Nils M. van Rooijen, Karlè V. Sýkora, Joop H.J. Schaminée

Abstract: The vegetation of coastal sand dunes is characterized by high species diversity and comprises some of the rarest vegetation types in North-Western Europe. Among them are dune grassland communities whose species richness relies on grazing. Those communities are assessed as a priority habitat type under the Natura 2000 legislation. In autumn 1990, Galloway cows and Nordic Fjord horses were introduced in the coastal dunes of Meijendel near The Hague (52°7'N, 4°20'E), The Netherlands, to reduce encroachment of tall grasses and shrubs, to develop bare sand patches, and as such facilitating diverse vegetation structures in the dune grasslands. In the 1950s, decades before the introduction of livestock, 41 permanent plots were installed. On average, they were examined every four years. Our study hypothesised that the livestock grazing in the set densities would halt progressive succession and facilitate regressive succession. Up to 1990, we observed an equilibrium between progressive and regressive succession. After 1990, however, our data showed a pronounced progressive succession contradicting the hypothesized effect of the livestock grazing. We relate the main observed patterns with two factors linked to rabbit populations: (i) the myxomatosis outbreak in 1954 and (ii) the rabbit Viral Haemorrhagic Disease (rVHD-1) outbreak in 1989. In addition to livestock grazing, rabbits block progressive succession by feeding on seedlings of shrub and tree species and digging burrows, creating small-scale mosaics of bare sand and initiate blowout development when collapsing. We state that the substantial decrease in rabbit numbers due to the viral diseases likely caused the observed increase of shrubs and trees in the study area's permanent plots. Climate change might have contributed to the observed increase in autonomous blowout development since 2001, as well as a decrease in atmospheric nitrogen deposition since 1990, after a strong increase the decades before.

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DOI: 10.3897/pls2022592/04

Authors: Michele Mugnai, Emilio Corti, Andrea Coppi, Daniele Viciani, Lorenzo Lazzaro

Abstract: We analysed two different plant communities hosting Ionopsidium savianum (Brassicaceae), a species of EU interest included in the Habitats Directive 92/43/CEE annexes, for which specific studies on the ecology of communities where the species grows are lacking and more in-depth knowledge is needed. We examined two important sites of occurrence of this species in Tuscany with different soil types, namely limestone (Mt. Calvi) and serpentine (Mt. Pelato), to determine the structural and functional profile of the communities hosting this species in such different contexts. At each site, we surveyed the plant communities with I. savianum in ten 1 m2 quadrats to determine information on communities' species composition and total plant cover, as well as taxonomic (species richness, and Shannon H’ index), phylogenetic (phylogenetic diversity, mean nearest taxon distance and mean pairwise distance) and functional diversity (focusing on Rao’s Q, leaf functional traits and adaptive strategies community weighted mean). We took into account site location, soil type, slope aspect and microrelief as plot-level environmental factors. The two communities were highly diverging from multiple points of view. Differences were in species composition, richness and diversity, with Mt. Calvi hosting higher diversity. The indices of phylogenetic diversity were influenced significantly by site and microrelief, allowing the presence of peculiar niches occupied by the fern Asplenium ceterach. From the functional point of view, communities at Mt. Calvi showed a higher functional diversity and a higher specific leaf area. Plant height was influenced by the slope aspect and was higher on north-facing slopes. In terms of Grime’s adaptive strategies, the Mt. Pelato communities resulted to be more stress tolerant than those surveyed at Mt. Calvi. Here, a decrease in stress-tolerant strategy associated with an increase in ruderal strategy was detected in communities on north-facing slopes.

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DOI: 10.3897/pls2022591/03

Authors: Daniele Viciani, Claudia Angiolini, Gianmaria Bonari, Alessandro Bottacci, Lorella Dell´Olmo, Vincenzo Gonnelli, Antonio Zoccola, Lorenzo Lastrucci

Abstract: The importance of wetlands for biodiversity conservation is widely recognized. Their relevance is crucial in the identification of habitat types included in the Habitats Directive 92/43/EEC and in the application of the following conservation measures. Yet, several montane and submontane areas are poorly investigated from the phytosociological point of view. Here we studied aquatic communities of the Tuscan-Romagna Apennines in Italy using 52 vegetation plots dominated by hydrophytes collected in wetlands of natural and artificial origin. We analysed our data using a cluster analysis and identified 12 vegetation types that we classified as three vegetation classes Charetea intermediae, Lemnetea and Potamogetonetea, and two habitat types of conservation importance, as the hard oligo-mesotrophic waters with benthic vegetation of Chara spp. (Natura 2000 code 3140) and the natural eutrophic lakes with Magnopotamion or Hydrocharition (Natura 2000 code 3150). Our study highlights the diversity of aquatic habitats in montane and submontane areas of the Northern Apennines where water bodies are generally small and scattered. This study further highlights the important role of natural freshwater ecosystems for vegetation, but also the notable role of partly or entirely artificial water bodies in maintaining aquatic communities.

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DOI: 10.3897/pls2021582/03

Authors: Benoît Vincent

Abstract: An opinion paper that tries to show that the concept of cenosis has been withheld over time, seeks to understand the reason why, and aims to rehabilitate it. Different definitions of biological and ecological communities types are then proposed. Finally, the paper presents some possible paths for a renewed science of cenosis (or cenology, or biocenotics).

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DOI: 10.3897/pls2021582/06

Authors: Mauro Fois, Gianluigi Bacchetta, Maria Carmela Caria, Donatella Cogoni, Emmanuele Farris, Giuseppe Fenu, Manuela Manca, Maria Silvia Pinna, Stefania Pisanu, Giovanni Rivieccio, Simonetta Bagella

Abstract: The ‘Habitats’ Directive (HD 92/43/EEC) is one of the primary legal tools aiming at conserving nature in Europe. Due to the complex iter to revise it, the habitats listed in the Annex I have been seldom updated after the HD adoption. Basing on already available information and expert knowledge, this paper presents a preliminary list of relevant habitats occurring in Sardinia, not yet considered and worth to be placed in the Annex I. Two new habitat proposals, one habitat new for Italy, and nine new subtypes of already existing HD habitats are here described. Most of the proposed new habitats and subtypes have a limited distribution range, due to the high number of narrow, often endangered, endemic species that characterize them. Being neglected, they are consequently poorly investigated, inconstantly monitored and unprotected. Thus, the main aim of this paper is to promote their conservation through implementation of HD and its interpretation manuals.

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DOI: 10.3897/pls2020581/08

Authors: Gianmaria Bonari, Edy Fantinato, Lorenzo Lazzaro, Marta Gaia Sperandii, Alicia Teresa Rosario Acosta, Marina Allegrezza, Silvia Assini, Marco Caccianiga, Valter Di Cecco, Annarita Frattaroli, Daniela Gigante, Giovanni Rivieccio, Giulio Tesei, Barbara Valle, Daniele Viciani, Giulia Albani Rocchetti, Claudia Angiolini, Emilio Badalamenti, Davide Barberis, Matteo Barcella, Giuseppe Bazan, Andrea Bertacchi, Rossano Bolpagni, Federica Bonini, Alessandro Bricca, Gabriella Buffa, Mariasole Calbi, Silvia Cannucci, Luigi Cao Pinna, Maria Carmela Caria, Emanuela Carli, Silvia Cascone, Mauro Casti, Bruno Enrico Leone Cerabolini, Riccardo Copiz, Maurizio Cutini, Leopoldo De Simone, Andrea De Toma, Michele Dalle Fratte, Luciano Di Martino, Romeo Di Pietro, Leonardo Filesi, Bruno Foggi, Paola Fortini, Roberto Gennaio, Gabriele Gheza, Michele Lonati, Andrea Mainetti, Marco Malavasi, Corrado Marcenò, Carla Micheli, Chiara Minuzzo, Michele Mugnai, Carmelo Maria Musarella, Francesca Napoleone, Ginevra Nota, Giovanna Piga, Marco Pittarello, Ilaria Pozzi, Safiya Praleskouskaya, Francesco Rota, Giacomo Santini, Simona Sarmati, Alberto Selvaggi, Giovanni Spampinato, Adriano Stinca, Francesco Pio Tozzi, Roberto Venanzoni, Mariacristina Villani, Katia Zanatta, Magda Zanzottera, Simonetta Bagella

Abstract: Habitat monitoring in Europe is regulated by Article 17 of the Habitats Directive, which suggests the use of typical species to habitat conservation status. Yet, the Directive uses the term “typical” species but does not provide a definition, either for its use in reporting or for its use in impact assessments. To address the issue, an online workshop was organized by the Italian Society for Vegetation Science (SISV) to shed light on the diversity of perspectives regarding the different concepts of typical species, and to discuss the possible implications for habitat monitoring. To this aim, we inquired 73 people with a very different degree of expertise in the field of vegetation science by means of a tailored survey composed of six questions. We analysed the data using Pearson’s Chi-squared test to verify that the answers diverged from a random distribution and checked the effect of the degree of experience of the surveyees on the results. We found that most of the surveyees agreed on the use of the phytosociological method for habitat monitoring and of the diagnostic and characteristic species to evaluate the structural and functional conservation status of habitats. With this contribution, we shed light on the meaning of “typical” species in the context of habitat monitoring.

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DOI: 10.3897/pls2020581/05

Authors: Miris Castello, Livio Poldini, Alfredo Altobelli

Abstract: Lake Doberdò (Classical Karst) is a well-known example of karst lakes, temporary lakes that seasonally fill and empty through springs and swallow holes connected to the underground waters. It is an area of exceptional interest for geological-geomorphological and biodiversity conservation, and is part of the Natura 2000 network. Its peculiar hydro-ecological nature allows an impressive variety of species and habitats. A phytosociological survey was carried out to provide a comprehensive analysis of the vegetation. On the basis of 177 phytosociological relevés and multivariate analysis 43 communities were found, belonging to Platyhypnidio-Fontinalietea antipyreticae, Lemnetea minoris, Potametea pectinati, Bidentetea tripartitae, Isoëto-Nanojuncetea, Phragmito-Magnocaricetea, Agrostietea stoloniferae, Molinio-Arrhenatheretea, Filipendulo ulmariae-Convolvuletea sepium, Alnetea glutinosae, Rhamno catharticae-Prunetea spinosae, Alno glutinosae-Populetea albae. The Leucojo aestivi-Poetum pratensis association including waterside periodically flooded meadows is validated. Compared to the past, various communities show a change in their distribution while some valuable coenoses were not found. 21 communities are attributed to 8 Annex I Habitats of the 92/43/EEC Directive. The study confirms the high value of Lake Doberdò for biodiversity conservation but highlights an ongoing process of environmental change due to both natural and human-related causes including modifications of the hydrological regime and abandonment of traditional agricultural practices. An overview of the features of Lake Doberdò is provided to compare this disappearing lake with the karst groundwater-dependent wetlands found in Ireland called “turloughs”, corresponding to Annex I Habitat 3180*. Also on the basis of the current interpretation and distribution of this habitat type at the EU level, a proposal is presented to recognize and protect this outstanding area of the Natura 2000 network in Italy as the habitat 3180*, modelled on Irish turloughs.

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DOI: 10.3897/pls2020572/06

Authors: Silvia Montecchiari, Marina Allegrezza, Veronica Pelliccia, Giulio Tesei

Abstract: Ailanthus altissima (tree of heaven), an invasive alien tree native to China, has become invasive all over the world and in Italy is present in all the administrative regions where it can form dense forest communities. Although there are several ecological studies on this species there is a lack of floristic-vegetational data for southern-Europe. The study presents the results of a floristic vegetational study on A. altissima forest communities of central Italy that aims to highlight the possible floristic-vegetational autonomy of these coenoses. The results have allowed the characterization of A. altissima coenoses at the ecological, biogeographic, syntaxonomic and landscape levels. These represent first A. altissima syntaxa described for the Italian peninsula and for southern-Europe. We propose two new sub-Mediterranean and Mediterranean associations comprised in the recently described alliance Lauro nobilis-Robinion pseudoaciae, in the Chelidonio-Robinietalia order and the Robinietea class: Asparago acutifolii-Ailanthetum altissimae: forest community with stratified structure and high canopy density on the warmer slopes of the hills in dry soil conditions and low anthropic disturbance and Aro italici-Ailanthetum altissimae: paucispecific forest communities with a monolayered structure typically found in agricultural, and peri-urban areas on pelitic, alluvial silty-sandy substrates, in conditions of edaphic humidity and high anthropogenic disturbance. The comparison with literature data highlights the autonomy of these associations of the sub-Mediterranean and Mediterranean alliance Lauro nobilis-Robinion pseudoacaciae alliance from the Balloto nigrae-Ailanthetum altissimae association of the Central and SE-European Balloto nigrae-Robinion pseudoacaciae alliance.

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DOI: 10.3897/pls2020572/03

Authors: Mattia Baltieri, Edy Fantinato, Silvia Del Vecchio, Gabriella Buffa

Abstract: Trait-based studies have become extremely common in plant ecology. In this work we analysed intraspecific trait variation of Himantoglossum adriaticum, a European endemic orchid species of Community interest, to investigate whether different populations growing on managed and abandoned semi-natural dry grasslands show differences in the CSR strategy. In seven populations occurring in Veneto Region (NE Italy), we measured H. adriaticum maximum vegetative height, leaf traits (LA, LDMC, SLA) and calculated the CSR strategy. Through CCA we investigated the relationship between plant traits and both plant community attributes (cover and height of herbs and shrubs), and geomorphologic features (aspects and slope). PERMANOVA test was used to investigate if the CSR strategy of H. adriaticum varied according to the management regime. Results showed that individuals of H. adriaticum develop different strategies when growing in different habitats. Specifically, individuals growing in managed fully sunny dry grasslands reached higher vegetative height (MH), had lower values of SLA and a higher relative contribution of the C parameter than individuals growing in abandoned dry grasslands, which, on the contrary, were shorter, had higher values of SLA (and correspondingly lower values of LDMC) and a higher relative contribution of the R parameter. Further data on reproductive traits (e.g. fruit and seed-set) may corroborate our results. Although the number of individuals addressed in this study is rather low, and our conclusions may not be considered of general validity for the species, our study demonstrated the applicability of the CSR strategy scheme in detecting functional strategies at intraspecific level.

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DOI: 10.3897/pls2020572/02

Authors: Lorenzo Gianguzzi, Giuseppe Bazan

Abstract: Knowledge of the processes by which plants colonize old structures is a key element for nature-based design both in urban and suburban contexts. This paper analyses the natural vegetation on walls and in other microhabitats of the roadway structures of Monte Pellegrino (606 m a.s.l.) near Palermo (Sicily), built in the first half of the 1900s. The historical road has particular construction and architectural features, and its characteristics have been maintained to this day. The route, approximately 16 kilometers long, is well integrated within a site of high naturalistic value which has been designated as a Special Area of Conservation (ITA020014) of the Natura 2000 network, and it is also a regional natural reserve. The survey was carried out on different homogeneous ecological contexts based on different microhabitats (masonry retaining walls, masonry guardwalls, road margins, and rock cut slopes) which are diversified according to other environmental factors (building materials, inclination, height, and exposure). The phytosociological and statistical analysis has led to the description of six new associations (Crepido bursifoliae-Parietarietum judaicae ass. nov., Athamanto siculae-Parietarietum judaicae ass. nov., Helichryso panormitani-Hypochaeridetum laevigatae ass. nov., Diantho siculi-Helichrysetum panormitani Gianguzzi ass. nov., Olopto miliacei-Pennisetetum setacei Gianguzzi ass. nov., Teucrio flavi-Rhoetum coriariae Gianguzzi ass. nov.) and one sub-association (Rhamno alaterni-Euphorbietum dendroidis Géhu & Biondi 1997 artemisietosum arborescentis subass. nov.). Other chasmophytic formations (Centranthetum rubri Oberd. 1969, Antirrhinetum siculi Bartolo & Brullo 1986) were reported for the first time in this area.

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DOI: 10.3897/pls2020571/09

Authors: Daniela Gigante, Gianni Bacchetta, Simonetta Bagella, Daniele Viciani

Abstract: The current issue is the first one of the new version of Plant Sociology, the international peer-reviewed journal of the "Società Italiana di Scienza della Vegetazione" (SISV). The technical management of the journal has been entrusted to the editorial platform Pensoft, the Editorial Board has been largely reshaped, now including also a dedicated Social media team. Plant Sociology is focused on all aspects of vegetation from phytocoenosis to landscape level, through time and space, at different geographic and ecological scales; the journal contributes to spread around the issues related to management and conservation of plant communities and plant diversity. All the articles are freely available in Open Access (OA) with affordable article processing charge (APC). In the present Editorial, we briefly discuss the importance of opening the access to knowledge and data about vegetation. We believe that disseminating plant science might be a precious tool for understanding ecological processes, modelling future trends and supporting decision makers. The introduced technological improvement will hopefully allow a larger visibility and circulation for the papers published on Plant Sociology.

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DOI: 10.3897/pls2020571/01

Authors: Marta Latini, Emanuele Fanfarillo, Elisa De Luca, Mauro Iberite, Giovanna Abbate

Abstract: The weed vegetation of the bean “Fagiolo Cannellino di Atina” (Phaseolus vulgaris L.) and the red pepper “Peperone di Pontecorvo” (Capsicum annuum L.) PDO (Protected Designation of Origin) crops was surveyed by means of 16 relevés, sampled in four farms of southern Latium during July 2019. The relevés were subjected to multivariate analysis, which revealed that the two crops are weeded by vegetation types referable to two different subassociations of Panico-Polygonetum persicariae (Spergulo-Erodion, Eragrostietalia, Digitario-Eragrostietea). Namely, communities colonizing bean fields, which are more mesophilous and richer in Eurasian taxa, are ascribable to the subassociation sorghetosum halepensis. Communities colonizing red pepper fields, which are more thermophilous and richer in Mediterranean taxa, are ascribable to the subassociation cyperetosum rotundi. Floristic, structural, and chorological features of the communities are discussed in relation to environmental factors and agricultural management.

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